Tyler Winklevoss

Winklevoss began rowing at the age of 15, encouraged by family friends and the example of a neighbor, Ethan Ayer, who rowed at Harvard University and Cambridge University. He began rowing at the Saugatuck Rowing Club on the Saugatuck River in 1997. His first coach was Irishman James Mangan who coached him and his brother throughout high school. Winklevoss' high school did not have a crew. In his junior year, he and his identical twin brother, Cameron Winklevoss, co-founded the crew program at their high school. In the summer of 1999, he made the United States Junior National Rowing Team, competing in the coxed pair event with his brother at the World Rowing Junior Championships in Plovdiv, Bulgaria.

He matriculated to Harvard College in 2000 and majored in economics, earning an A.B. and graduating in 2004. At Harvard, he was a member of the men's varsity crew, the Porcellian Club and the Hasty Pudding Club.

In December 2002, Winklevoss, along with his brother Cameron Winklevoss and fellow Harvard classmate Divya Narendra, sought a better way to connect with fellow students at Harvard University and other universities. The three conceived of a social network for Harvard students named HarvardConnection; the concept ultimately expanded to other schools around the country. In January 2003, they enlisted the help of fellow Harvard student, programmer and friend Sanjay Mavinkurve to begin building HarvardConnection. Mavinkurve commenced work on HarvardConnection but departed the project in spring 2003 when he graduated and went to work for Google.

In November 2003, at the suggestion of Victor Gao, the Winklevosses and Narendra approached Mark Zuckerberg about joining the HarvardConnection team. The previous HarvardConnection programmers had reportedly made progress on coding front-end pages, the registration system, a database, back-end coding, and a way users could connect with each other, which Gao called a "handshake". In early November, Narendra emailed Zuckerberg saying, "We're very deep into developing a site which we would like you to be a part of and ... which we know will make some waves on campus." Within days, Zuckerberg was talking to the HarvardConnection team and preparing to take over programming duties from Gao. On the evening of November 25, 2003, the Winklevosses and Narendra met with Zuckerberg in the dining hall of Harvard's Kirkland House, where they explained to Zuckerberg the HarvardConnection website, the plan to expand to other schools after launch, the confidential nature of the project, and the importance of getting there first. During the meeting, Zuckerberg allegedly entered into an oral contract with Narendra and the Winklevosses and became a partner in HarvardConnection. He was given the private server location and password for the unfinished HarvardConnection website and code, with the understanding that he would finish the programming necessary for launch. Zuckerberg allegedly chose to be compensated through an interest in the enterprise (sweat equity).

On November 30, 2003, Zuckerberg told Cameron Winklevoss in an email that he did not expect completion of the project to be difficult. Zuckerberg writes: "I read over all the stuff you sent and it seems like it shouldn't take too long to implement, so we can talk about that after I get all the basic functionality up tomorrow night." The next day, on December 1, 2003, Zuckerberg sent another email to the HarvardConnection team. "I put together one of the two registration pages so I have everything working on my system now. I'll keep you posted as I patch stuff up and it starts to become completely functional." On December 4, 2003, Zuckerberg writes: "Sorry I was unreachable tonight. I just got about three of your missed calls. I was working on a problem set." On December 10, 2003: "The week has been pretty busy thus far, so I haven't gotten a chance to do much work on the site or even think about it really, so I think it's probably best to postpone meeting until we have more to discuss. I'm also really busy tomorrow so I don't think I'd be able to meet then anyway." A week later: "Sorry I have not been reachable for the past few days. I've basically been in the lab the whole time working on a cs problem set which I'm still not finished with." On December 17, 2003, Zuckerberg met with the Winklevosses and Narendra in his dorm room, allegedly confirming his interest and assuring them that the site was almost complete. On the whiteboard in his room, Zuckerberg allegedly had scrawled multiple lines of code under the heading "Harvard Connection," however, this would be the only time they saw any of his work. On January 8, 2004, Zuckerberg emailed to say he was "completely swamped with work [that] week" but had "made some of the changes ... and they seem[ed] to be working great" on his computer. He said he could discuss the site starting the following Tuesday, on January 13, 2004. On January 11, 2004, Zuckerberg registered the domain name thefacebook.com. On January 12, 2004, Zuckerberg e-mailed Eduardo Saverin, saying that the Facebook site [thefacebook.com] was almost complete and that they should discuss marketing strategies. Two days later, on January 14, 2004, Zuckerberg met again with the HarvardConnection team; however, he allegedly failed to disclose registering the domain name thefacebook.com or developing a competing social networking website. Rather, he allegedly reported progress on HarvardConnection, told the team he would continue to work on it, and would email the group later in the week. On February 4, 2004, Zuckerberg launched thefacebook.com, a social network for Harvard students, designed to expand to other schools around the country.

On February 6, 2004, the Winklevosses and Narendra first learned of thefacebook.com while reading a press release in the Harvard student newspaper The Harvard Crimson. According to Gao, who looked at the HarvardConnection code afterward, Zuckerberg had left the HarvardConnection code incomplete and non-functional, with a registration that did not connect with the back-end connections. On February 10, 2004, the Winklevosses and Narendra sent Zuckerberg a cease and desist letter. They also lodged a complaint with the Harvard administration regarding what they viewed as a violation of the university's honor code and student handbook. The Harvard Administrative Board and university president Larry Summers reportedly viewed the matter to be outside of the university's jurisdiction. President Summers advised the HarvardConnection team to take their matter to the courts.

In 2004, ConnectU filed a lawsuit against Facebook alleging that creator Mark Zuckerberg had broken an oral contract with it. The suit alleged that Zuckerberg had copied ConnectU's idea and illegally used source code intended for the website Zuckerberg was hired to develop. Facebook countersued with respect to Social Butterfly, a Winklevoss Chang Group project. The countersuit named among the defendants ConnectU, Cameron Winklevoss, Tyler Winklevoss, Divya Narendra, and Wayne Chang, founder of i2hub. An agreement settling both cases was reached in February, 2008, with the Winklevoss party receiving $20 million in cash and $45 million in Facebook stock. In May 2010, however, ConnectU accused Facebook of misrepresenting the value of the stock that it turned over to the ConnectU plaintiffs as part of the settlement and sought to void the settlement. ConnectU alleged that the value of the stock was $11 million rather than $45 million, as represented by Facebook at the time of settlement. As a result, the total settlement value would have been $31 million, rather than the $65 million reported. On August 26, 2010, the New York Times reported that Facebook shares were then trading at $76 per share in the secondary market, putting the value of the total settlement at close to $120 million. If the lawsuit to revise the settlement were to succeed, the settlement value would rise to $466 million. In April, 2011, Ninth Circuit judge Alex Kozinski opined that "[a]t some point, litigation must come to an end. ... That point has now been reached." The twins' lawyer said they will seek a rehearing with the entire appeals court bench. In June 2011 it was announced that a decision to pursue the case in the Supreme Court had been withdrawn.

Winklevoss rowed at Harvard University for 4 years under coach Harry Parker, while completing his undergraduate studies. In 2004, he sat 5-seat in the "engine room" of the Harvard men's varsity heavyweight eight boat. The 2004 crew was nicknamed the "God Squad" because, according to Winklevoss, some of them believed in God while the rest believed they were god. As a Harvard Crimson in 2004, they thought he helped the "God Squad" win the Eastern Sprints, the Intercollegiate Rowing Association Championship, and the Harvard–Yale Regatta to cap-off an undefeated collegiate racing season.

In 2007, Winklevoss was named to the United States Pan American Team and competed at the 2007 Pan American Games in Rio de Janeiro, Brazil. He won a silver medal in the men's coxless four event and stroked the men's eight boat to a gold medal on the Lagoa Rodrigo de Freitas.

In 2008, Winklevoss was named to the United States Olympic Team and competed at the 2008 Olympic Games in Beijing, China. He rowed with his brother in the men's coxless pair event which took place at the Shunyi Olympic Rowing-Canoeing Park. The brothers were coached by the renowned Ted Nash. In their first heat, they failed to finish in the top three and did not qualify for the Semifinals. In the Repechage (a last chance to make the Semifinals), they took first, advancing them to the Semifinals. A strong finish in Semifinal 2 put them in the Final. They ended up finishing sixth out of the fourteen countries which had qualified for the Olympics.

In 2009, Winklevoss began graduate business study at the Saïd Business School at the University of Oxford and completed an MBA in 2010. While at Oxford, he was a member of Christ Church, an Oxford Blue, and rowed in the Blue Boat in the Oxford-Cambridge Boat Race.

On December 21, 2009, i2hub founder Wayne Chang and The i2hub Organization launched a lawsuit against ConnectU and its founders, Cameron Winklevoss, Tyler Winklevoss, and Divya Narendra, seeking 50% of the settlement proceeds from the original lawsuit. The complaint says "The Winklevosses and Howard Winklevoss filed [a] patent application, U.S. Patent Application No 20060212395, on or around March 15, 2005, but did not list Chang as a co-inventor. It also states "Through this litigation, Chang asserts his ownership interest in The Winklevoss Chang Group and ConnectU, including the settlement proceeds." Lee Gesmer of Gesmer Updegrove, LLP posted the detailed 33-page complaint online.

Between November 30, 2003, and February 4, 2004, Zuckerberg exchanged a total of 52 emails with the Harvard Connection team and engaged in multiple in-person meetings. During the same period of time, Zuckerberg engaged in multiple electronic instant message communications with people outside of the HarvardConnection team. On March 5, 2010, certain electronic instant messages from Mark Zuckerberg's hard drive were leaked to the public. On September 20, 2010, Facebook confirmed the authenticity of these leaked instant messages in a New Yorker article.

One of ConnectU's law firms, Quinn Emanuel, inadvertently disclosed the confidential settlement amount in marketing material by printing "WON $65 million settlement against Facebook". Quinn Emanuel sought $13 million as its contingency fee related to the original settlement. ConnectU fired Quinn Emanuel and sued the law firm for malpractice. On August 25, 2010, an arbitration panel ruled that Quinn Emanuel "earned its full contingency fee." The panel also found that Quinn Emanuel committed no malpractice.

On May 13, 2011, it was reported that Judge Peter Lauriat made a ruling against the Winklevosses. Chang's case against them could proceed. The Winklevosses had argued that the court lacks jurisdiction because the settlement with Facebook has not been distributed and therefore Chang hasn't suffered any injury. Judge Lauriat wrote, "The flaw in this argument is that defendants appear to conflate loss of the settlement proceed with loss of rights. Chang alleges that he has received nothing in return for the substantial benefits he provided to ConnectU, including the value of his work, as well as i2hub's users and goodwill." Lauriat also wrote that, although Chang's claims to the settlement are "too speculative to confer standing, his claims with respect to an ownership in ConnectU are not. They constitute an injury separate and distinct from his possible share of the settlement proceeds. The court concludes that Chang has pled sufficient facts to confer standing with respect to his claims against the Winklevoss defendants."

In 2012, Tyler and his brother Cameron founded Winklevoss Capital Management, a firm that invests across multiple asset classes with an emphasis on providing seed funding and infrastructure to early-stage startups. The company is headquartered in New York's Flatiron District.

The twins were depicted on the animated television show The Simpsons in the eleventh episode of Season 23 in the episode called "The D'oh-cial Network" which aired on January 15, 2012. The Winklevoss twins are seen rowing in the 2012 Olympic Games against Marge Simpson's sisters Patty and Selma. There is a reference made to the $65 million Facebook settlement.

In 2014, Tyler and his brother Cameron founded Gemini, a digital currency exchange and custodian that allows customers to buy, sell, and store digital assets. It is a New York trust company that is regulated by the New York State Department of Financial Services (NYDFS).